If I were you, I would try many combinations to find out.First try PCI-E #1 and #3 and compare findings with these you have now.
This is not the answer you were looking for, but it could be helpful.
Anyway, look down to my reply to kpablo and what I'm going to do, so we'll know if #2 and #4 run @16x if #1 and #3 are not populated.
I doubt that, but not sure. I'll try putting my GTX 480 from slot 1 to slot 2 and post result.
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